Protocol to Quantify Nematophagous Fungi in Soil

This is an outline and modification of methods developed by Jaffee et al. (1996):

1. Prepare quarter-strength cornmeal agar (CMA/4)1 amended with 0.1 g/L of streptomycine sulfate after the CMA/4 is cooled down to approximately 50°C (does not burn your hand). Pour the agar medium into 9-cm-diameter or 6-cm-diameter petri dishes. Prepare 15 plates of CMA/4 for each sample.

2. Subsample 100 g of soil from each sample, placed in a 250-mL beaker, and add 200 mL sterile distilled water. Stir the suspension vigorously for 3 minutes.

3. Conduct 10-fold dilution of the well-mixed soil suspension twice. This will provide three soil dilutions: 0.5, 0.05, and 0.005g of soil/mL H2O.

4. Each dilution will be plated on 5 replicated petri dishes. Each petri dish will receive 100 ml of soil dilutions. Thus, a total of 15 plates for each sample with 5 plates each of 0.05, 0.005, and 0.0005 g of soil / plate.

5. Add approximately 1000 Steinernema glaseri (a large and long-lived entomophilic nematode) in 0.1 mL distilled water to each plate as a bait for nematode-trapping fungi. This will increase the chances of detecting nematode-trapping fungi.

6. Infective juveniles of S. glaseri can be obtained from infected Galleria mellonella  according to protocol by Woodring and Kaya (1988) and stored at 10°C.

7. Observe the entire surface of each petri dish 3 weeks after plating under a dissecting microscope with 50-140 ´ magnification.

8. Record the presence and absence of nematode-trapping fungi in species using the Cooke and Godfrey key (1964) for each petri dish.

9. Use a most-probable number table (Alexander, 1982) or see the Most Probable Number Table below to estimate the number of fungal propagules per g soil.

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1 Prepare CMA/4 by mixing quarter strength of Corn Meal Agar (Becton Dickinson and Company, Sparks, MD), i.e. 4.25 g/L H2O with three quarter strength of Bacto Agar (Difco Laboratories, Detroit, MI) i.e. 8.25 g/L H2O.


Most Probable Number (MPN) Table

 

Base of dilution ratio = 10

No plates/dilution = 5

Inoculation vol. (in mL) =  0.1

Initial dilution is 1 to = 2

 

 

No. of positive plates on dilutions

 

 

 

0.05

0.005

0.0005

MPN

P(0.95)

0

0

0

0

-

0

1

0

3.6328

1.1002-11.9952

1

0

0

3.9690

1.2020-13.1053

1

1

0

8.0454

2.4366-26.5653

2

0

0

8.9257

2.7032-29.4719

2

1

0

13.6612

4.1373-45.1079

3

0

0

15.5307

4.7035-51.2812

3

1

0

21.3767

6.4740-70.5844

3

2

0

27.6370

8.3699-91.2553

4

0

0

25.4592

7.7104-84.0646

4

1

0

33.6994

10.2060-111.2728

4

2

0

43.1111

     13.0564-142.3500

4

3

0

54.0130

     16.3580-178.3470

5

1

0

65.4321

     19.8163-216.0520

5

2

0

97.7203

     29.5949-322.6654

5

3

0

156.1747

     47.2980-515.6776

5

4

0

254.5840

     77.1016-840.6184