Koon-Hui
Wang, Robert McSorley
University of Florida, Department of
Entomology and Nematology, P.O. Box 110620, Gainesville, FL 32611-0620, U.S.A.
(Last updated September 2003)

A key to commonly
occurring nematophagous fungi
Ecology of nematophagous fungi
Protocol to
quantify nematophagous fungi
Examples of use of cover crop
for enhancement of nematophagous fungi
Factors affecting
cover crops for nematophagous fungi
Duration of cover crop for
enhancing nematophagous fungi in field conditions
Nematophagous
fungi are fungi that feed on nematodes. These fungi can be fungal
egg-parasites, nematode-trapping fungi that capture nematodes using modified
hyphal traps, or endoparasites that parasitize the nematode by means of small
conidia or zoospores. There are various ways for soil-borne fungi to suppress
nematode multiplication. A detailed review of fungi as biocontrol agents
against plant-parasitic nematodes has been published (Kerry and Jaffee, 1997). In summary, there are five mechanisms that fungi
use to suppress nematodes. Some of these interactions are direct whereas others
are indirect. The direct mechanism is performed by 1) fungi that feed on
nematodes directly, known as nematophagous fungi; fungi interact with nematodes
in an indirect manner by several mechanisms including: 2) fungi that kill
nematodes by mycotoxin (Barron and
Thorne, 1987) or 3) through the destruction of the feeding sites
of sedentary nematodes in roots (Glawe
and Stiles, 1989); 4) fungi that are nonpathogenic to plants, but
compete with nematodes in roots and significantly reduce nematode
multiplication (Sikora, 1992). Many of these fungi are used as potential nematode
biocontrol agents; 5) Mycorrhizal fungi improve the growth of nematode infected
plants and may also affect nematode development (Hussey and Roncadori, 1982).
According to a survey of nematophagous fungi in
Ireland by Gray (1983), nematophagous fungi were found in all of the habitats
examined, among which, permanent pasture, coniferous leaf litter, and coastal
vegetation had the most frequent incidence of nematophagous fungi. Other
habitats examined by Gray included coniferous leaf litter, old and partly
revegetated dung, permanent grassland pasture, cultivated land, moss cushions,
decaying vegetation and compost, and peatland (Gray, 1983). In addition, many
other studies (Barron, 1977) also supported the hypothesis that nematophagous
fungi are widely distributed and have great potential to be explored as
biocontrol agents.
However, as
stated by Kerry (1988): “The successful introduction of such an
agent depends on whether a suitable niche for the microorganism exists or can
be created and until we know much more about the factors that affect the
activity of nematophagous fungi in soil, their full potential as control agents
for nematodes will not be realized.”
This article
summarizes groups of nematophagous fungi according to their feeding habits:
A) Nematode-trapping fungi
Facultative fungi that form trapping structure to trap nematodes. There are 6 types of traps reported by Barron (1977).
adhesive hyphae:
eg. Zygomycotina
Stylopage
Cystopage
adhesive traps:
eg. Deuteromycota
Monacrosporium
cionopagum (branches)
M.
ellipsosporum (knobs)
Arthrobotrys
oligospora (networks)
non-adhesive traps
eg. Deuteromycota
Arthrobotrys
dactyloides (constricting ring)
Dactylella leptospora (non-constricting ring)
B) Facultative parasitic fungi attacking
sedentary stages of nematodes (Kerry and Jaffee, 1997)
These
are facultative fungi that are commonly soil saprophytes, and are opportunistic
fungi isolated from the sedentary stages (female and egg stages) of sedentary
nematodes such as Heterodera, Globodera, and Meloidogyne. They do
not form specialized infection structures except appressoria. They can survive
and proliferate in soil in the absence of nematodes.
eg.
Hyphomycotina
Acremonium
Cylindrocarpon
Fusarium
Paecilomyces
Verticillium
C) Endoparasitic fungi (Kerry and Jaffee,
1997)
These are obligate parasitic fungi that have limited growth in soil outside the colonized nematode cadaver.
They can infect vermiform nematodes by
producing adhesive spores attached to cuticle of passing nematodes.
eg. Hyphomycotina
Hirsutella
rhossiliensis
Drechmeria coniospora
Verticillium spp.
Some can infect vermiform nematodes by
producing conidia spores that can be ingested by nematodes.
Harposporium
anguillulae
Some can infect vermiform nematodes by
producing motile zoospores that encyst on the nematode’s surface.
eg. Oomycota
Myzocytium spp.
Lagenidium spp.
Chytridiomycota
Catenaria anguillulae
Some can infect sedentary nematodes when the
nematodes were exposed on the root surface.
eg. Oomycota
Nematophthora gynophila
An
ecology study of nematophagous fungi conducted by Gray (1985) revealed that
different types of nematophagous fungi have different edaphic preferences. The
saprophytic NTF (formed adhesive nets) are found in soil with low organic
matter and low moisture due to their saprophytic nature. When nutrients or
moisture condition improved, the saprophytic NTF are able to compete with other
soil organisms by feeding on the expanding nematode population. In contrast,
NTF that form rings are more common in soil with high organic matter and
moisture. Endoparasitic fungi that produce conidia are strongly influenced by
organic matter. While most of the NTF (except those that formed adhesive
branches) are not affected by nematode densities, endoparasitic fungi that form
ingestive spores are nematode-density dependent. In general, the
conidia-forming endoparasites were isolated from samples with comparatively
high soil moisture and low pH. Little is known about edaphic preference of the
nematophagous fungi with unmodified adhesive hyphae, except that they are more
frequently recovered from soils with higher pH. Table 1-1 summarizes soil
factors affecting different nematophagous fungi.
Table 1. Effect of soil edaphic factors on distribution of nematophagous fungi (Gray, 1985).
|
Nematophagous fungi |
Organic matter |
pH |
Moisture |
Nematode densities |
|
Nematode-trapping |
NSz |
low |
NS |
NS |
|
net |
lowy |
low |
low |
NS |
|
ring |
highx |
low |
high |
NS |
|
adhesive hyphae |
NS |
high |
NS |
NS |
|
adhesive branch |
NS |
NS |
NS |
high |
|
adhesive knobs |
NS |
low |
NS |
NS |
|
Endoparasites |
high |
low |
high |
high |
zNS = effects of the
edaphic factors is not significant.
ylow = lower value of the
edaphic factor is preferred by the group of nematophagous fungi.
xhigh = higher value of
the edaphic factor is preferred by the group of nematophagous fungi.
Base on their
ecological preferences, nematode-trapping fungi (NTF) are separated into two
groups: saprophytic and parasitic NTF (Cooke,
1963).
Saprophytic
NTF—form 3-dimensional-network traps in response to the presence of nematodes.
Under low nematode population densities, they remain saprophytic. Therefore,
they are regarded as inefficient nematode-trappers.
Parasitic
NTF—have low saprophytic ability, but form traps spontaneously. This group
consists of NTF that form constricting rings, adhesive branches and are more
effective nematode trappers than the saprophytic NTF (Jasson, 1982).
Further
information on nematode-antagonistic fungi can be obtained through:
http://sacs.cpes.peachnet.edu/nemabc/
http://www.area.ba.cnr.it/~e085ac01/bkfair3444.html
http://www.ag.auburn.edu/bci/nematolo.htm
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